From my little knowledge, the ad networks place ads on your site and get to pay the site owner for the number of views for ads displayed to users if the site.
Advertising networks connects the website owners with advertisers. In order to use them to generate income one must first of all sign up with good ad networks.
A business wants to promote something, it runs ad campaign through ad network. The ad network runs the ads on partner website. The publishers earn the share of advertising budget
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